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The utricle and saccule are the two otolith organs in the vertebrate inner ear. They are part of the balancing system (membranous labyrinth) in the vestibule of the bony labyrinth (small oval chamber). They use small stones and a viscous fluid to stimulate hair cells to detect motion and orientation. The utricle detects linear accelerations and head-tilts in the horizontal plane. The word utricle comes from Latin uter 'leather bag'.